I am an Emeritus Professor of Biology at Case Western Reserve University.  I received my A.B. Degree from Dartmouth College and M.S. and Ph.D. Degrees from the University of Wisconsin, Madison, in Zoology.  I joined the faculty of the Department of Biology at Case Western Reserve University in 1973 and retired as a Professor of Biology with a secondary appointment as Professor in the Department of Electrical Engineering and Computer and Science in July 2012.  I served as Chair of the Department of Biology at Case Western Reserve University from 2000 to 2009. 
I have broad research interests in aquatic systems ecology and have served on a number of Boards and advisory committees for the Great Lakes Fishery Commission, the International Joint Commission, National Research Council, and the U.S. Environmental Protection Agency.  My experience in undergraduate biology teaching and curriculum development includes service as a Program Director for CWRU’s Howard Hughes Medical Institute grant and recognition as a National Academies Education Fellow in the Life Sciences in 2004-2005, and I was elected a SENCER Leadership Fellow in 2012. I served as a co-Director of the SENCER Center for Innovation—Great Lakes from 2012 to 2015.

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